It was a hug worthy only of the woman who redefined man.

For Jane Goodall, that 2014 embrace with a rescued chimp called Wounda at her institute's sanctuary in the Republic of Congo was yet another vindication of her life's work.

Because more than 50 years earlier she had become the first person to document any animal other than humans using a tool, when she observed a chimp digging termites out of a mound with a stick.

Dr Goodall, who has died aged 91, had proven in that moment in Tanzania in 1960 the indelible link between modern man and our ape cousins.

As her great mentor, the palaeoanthropologist Louis Leakey, had said in response to her discovery: 'Now we must redefine tool, redefine man, or accept chimpanzees as human.'