World-renowned chimpanzee expert Jane Goodall died of cardiopulmonary arrest, according to the death certificate obtained by TMZ.

The primatologist was 91 when she died in Los Angeles on Oct. 1, while she was in the U.S. on a speaking tour.

Goodall’s death certificate also indicated she had epilepsy, though it is not clear whether that factored into her death.

The Jane Goodall Institute said its founder died “due to natural causes” when it announced her death earlier this month.

In the statement, the organization said, “Dr. Goodall’s discoveries as an ethologist revolutionized science, and she was a tireless advocate for the protection and restoration of our natural world.”